Biological cultural heritage may refer to a site where historical management has created species-rich grasslands, as well as to single species. a A semi-natural grassland at the Natura 2000 site of Stora Åsa, Province of Södermanland, Sweden. b Succisa pratensis, a species characteristic for these grasslands and which remains as legacy in forests long after management has ceased (Herben et al. 2006). Photo: Ove Eriksson.